If a different monkey, who had not participated in the game, was given the prize of grapes instead of the female participating in a bid to win them, the monkeys demonstrated aggression towards the experimenter. In a later study, Brosnan et al discovered that chimps were more upset by an injustice in a casual relationship with another primate than in a more intimate relationship. The results of these studies are important since they suggest that explanations for romantic relationships, including equity theory, have an evolutionary basis.