during major activity, the muscle cells around the tracheoles carry out anaerobic respiration. this produces lactate, which is soluble and lowers the water potential of muscle cells. so water moves into cells from the tracheoles by osmosis. the water in the ends of the tracheoles decreases in volume and so draws air further into them. this means the final diffusion pathway is in a gas rather than liquid phase. so diffusion is more rapid. this increases the rate that air is moved into the tracheoles but leads to more water evaporation