Mouth opens (operculum is closed), buccalcavity floor is lowered, increasing volume and decreases the pressure of the buccal cavity compared to the outside so water rushes in the mouth down a pressure gradient
2. Opercular cavity expands, buccalcavity is raised, pressure inside the buccalcavity is now higher than in the opercular cavity so water moves from the buccalcavity over the gills into the opercular cavity
3. The mouth is now closed and the operculumopens, sides of the opercular cavity move inwards, increasing in pressure and water rushes out of the fish through the operculum