Biological evolution: Origin of life, diversification into novel forms (species), change in genetic composition of populations over generation, in the past and into the future.
Descent with modification -> understanding of the changes in the genetic composition of populations and the processes underlying those changes. Investigation of origins and distribution of species.
Study evolution -> to clarify the origin and history of populations and species, patterns of diversification and geographic distribution
Understand genetic changes that occurred in the past and ongoing genetic changes into the future.
Genetic change of populations result from multiple distinct process: Recombination (mating patterns), Mutation, Selection, Random genetic sampling 'genetic drift', Inter-population movement 'gene flow'.
Over extended time periods of time, these processes can result in divergence among populations and species.
Plato, Aristotle -> fixity of organisms (biodiversity)
Uniformitarianism: change is gradual, with continuity; natural laws are consistent.
Scientists propose theories based on observations (evidence) from the natural world, and following their reasoning towards logical conclusions.
Adaptive evolution: evolution that increases the suitability of a population to the habitat it occupies and consequently, its chance to continue persisting there.
Darwin and Wallace made extensive collections and observations documenting extensive variation within species, as well as among them.
Variation is, in part, heritable.
Heritability of differences: for many traits that vary within a population, individuals appear more similar to closer relatives; in particular, offspring resemble their parents.
Selective breeding takes advantage of heritable variation.
Descendant species resemble each other because they inherit their characteristic traits from a common ancestor.
Evidence of common ancestry in nature -> Biogeography - the geographical distribution of biodiversity reflects common ancestry
Of all juveniles produced, few survive to produce offspring : resources do not suffice for all; many die before reproducing.
Variation in survival and reproduction, natural selection, could result from resource limitation.
C&C: Lamarck's hypothesis: Transmission parent to offspring of acquired characteristics --> believed in spontaneous generation as ongoing process
C&C: Darwin's hypothesis: Transmission parent to offspring of inherent trait differences --> Formulated idea of "descent with modification"
Disproving spontaneous generation was key to the development of evolutionary theory.
Life arises from pre-existing life, not spontaneously from non-living materials.
Obs: Individuals within species vary in phenotype.
Obs: Offspring look like their parents. Phenotypic differences are, to some extent, heritable, transmitted from parent to offspring.
Obs: Individuals differ in their survival and reproduction, they differ in fitness.
If survival and reproduction depend on heritable phenotypic differences then evolution by natural selection (adaptive evolution)
Species comprise individuals that vary ever so slightly from each other with respect to their many traits.
Species have a tendency to increase in numbers over generations at a geometric rate.
Natural selection and artificial selection follow the same principles. Distinction : Natural mediated by Environment, Artificial mediated by human choices.
Selection: some individuals, because of their traits relative to their environment, are more likely to survive and reproduce than others. They have a higher fitness.
Higher fitness -> offspring are likely to have inherited values similar to those of their parents. These offspring represent a distinctive subset of their parents. The overall trait dist. of the population and its genetic composition differ from that of the population in the previous generation.
Natural selection depends on heritable variation of Zinc tolerance in grass.
Selection: a process occurring within a generation: differential survival and reproduction. Results in change in a population's genetic composition, but the only one that directly promotes increase in average fitness.
Adaptation: a trait that contributes to fitness and that has been honed by selection. A multigenerational process, mediated by natural selection, by which a population's average fitness in its environment increases.
Evolution does not progress from "less" to "more" advanced. It is not goal-seeking.
'Survival of the fittest': overemphasizes variation in survival, competition/struggle, an extreme simplification of evolution.
Over larger time scales, Darwin understood that descendant species inherited traits from their ancestral species.
Particulate inheritance: characteristics of both parents are passed on, but not necessarily visible in phenotypes. (Maintains variation!)
Law of segregation: each gamete is haploid; carries one allele per genetic locus
Law of independent assortment: alleles at different loci/chromosomes segregate independently of one another --> new combination of alleles