Pioneer species (first species to colonise an area)
Colonise bare rock
Seeds/Spores blown in by wind + grow
Harsh abiotic conditions (soil has low H2O retention)
Pioneer species specially adapted
They change the abiotic conditions less hostile (break rock, die, add dead organic matter (humus) as they decompose)
Now soil can retain H2O
So new organisms can colonise + die
Increasing organic material of soil
New species change environment
Less suitable for previous species
Previous species outcompeted by larger plants