mouth opens and buccal cavity is lowered
volume of buccal cavity increases, and pressure decreases so water enters
simultaneously, the opercular valve is shut and the opercular cavity containing gills expands
pressure in the opercular cavity decreases
floor of the buccal cavity move upwards, increasing pressure - water then moves from buccal cavity over the gills
mouth closes and operculum opens, sides of opercular cavity move inwards, increasing pressure, forcing the water over the gills and out the operculum
buccal cavity moves up slowly to maintain the flow of water