Buccal cavity drops and mouth opens increasing volume of buccal cavity and decreasing pressure so that water flows into the buccal cavity down a pressure gradient. Simultaneously, operculum valve shuts and operculum cavity expands. Volume of the operculum cavity increases and decreases pressure. Fish close mouth and buccal cavity raises, decreasing volume increasing pressure inside the buccal cavity, water in buccal cavity moves over gills. Operculum opens, decreasing pressure in operculum cavity and water moves out the opercular opening.