Photosystems use some wavelengths of light but reflect others
Photosynthetic pigments
Arranged as "photosystems"
Chlorophyll
A pigment molecule that functions to capture (absorb) light energy
Chlorophyll doesn't capture light energy alone, it works with several hundred chlorophyll molecules and other pigments like carotenoids or xanthophylls
Photosystems use some wavelengths of light but reflect others
Light is trapped and absorbed by a network of chlorophyll pigments
Chlorophyll
A pigment molecule that functions to capture (absorb) light energy
Chlorophyll doesn't capture light energy alone, it works with several hundred chlorophyll molecules and other pigments like carotenoids or xanthophylls
Photosystems
1. Chlorophyll molecules absorb photons and raise their electrons to a higher energy level
2. Excited electrons reduce a reaction center, the primary acceptor in a redox reaction
3. This process is called photo-oxidation
Light is trapped and absorbed by a network of chlorophyll pigments
There exists 2 photosystems that function to gather light energy
Photosystems
1. Chlorophyll molecules absorb photons and raise their electrons to a higher energy level
2. Excited electrons reduce a reaction center, the primary acceptor in a redox reaction
3. This process is called photo-oxidation
Robert Emerson determined that efficiency of photosynthesis greatly increased when he combined short and long wavelengths, concluding that two photosystems existed and photosynthesis worked best when both were operating
There exists 2 photosystems that function to gather light energy
Photosystem I and photosystem II
The two light-capturing complexes used by most photoautotrophs
Robert Emerson determined that efficiency of photosynthesis greatly increased when he combined short and long wavelengths, concluding that two photosystems existed and photosynthesis worked best when both were operating
Photosystem I and photosystem II
The two light-capturing complexes used by most photoautotrophs
The electron carriers of the photosynthetic system consist of non-protein organic groups that alternate between being oxidized and being reduced as electrons move through the system
The carriers include compounds that are similar in structure and function to those in mitochondrial electron transport chains
The electron carriers of the photosynthetic system consist of non-protein organic groups that alternate between being oxidized and being reduced as electrons move through the system
Light Harvesting Pigments
Absorb light energy and transfer it to the Reaction Center
The carriers include compounds that are similar in structure and function to those in mitochondrial electron transport chains
Reaction Center
Where the absorbed light energy is used to drive electron transport
Photosystems
Light Harvesting Pigments
Reaction Center
Photon
Photon
A particle of light
Photosynthetic Electron Transport Chain (PETC)
Electrons are passed along from the Photosystem
Photosystem
The light-capturing complexes that contain the light harvesting pigments and reaction centers
Light Reactions
1. Oxidation of P680
2. Oxidation-reduction of plastoquinone
3. Electron transfer from the cytochrome complex and shuttling by plastocyanin
4. Oxidation-reduction of P700
5. Electron transfer to NADP+ by ferredoxin
6. Formation of NADPH
Photosynthetic Electron Transport Chain (PETC)
Electrons are passed along from the Photosystem
The proton gradient is used to generate ATP with the same kind of ATP synthase complexes found in mitochondrial membranes
Light Reactions
1. Oxidation of P680
2. Oxidation-reduction of plastoquinone
3. Electron transfer from the cytochrome complex and shuttling by plastocyanin
4. Oxidation-reduction of P700
5. Electron transfer to NADP+ by ferredoxin
6. Formation of NADPH
Chemiosmotic Synthesis of ATP
1. Protons are taken into the lumen by the reduction and oxidation of plastoquinone
2. The concentration of protons inside the lumen is increased by the addition of two protons for each water molecule that is split in the lumen
3. The removal of one proton from the stroma for each NADPH molecule formed decreases the concentration of protons in the stroma outside the thylakoid
The absorption of light energy by photosystem II results in the formation of an excited-state P680 (P680*) molecule, which is rapidly oxidized, transferring a high-energy electron to the primary acceptor
The higher concentration of protons inside the membrane creates a substantial proton-motive force that drives protons out of the lumen, back into the stroma
Oxidation-reduction of plastoquinone (PQ)
1. PQ accepts electrons from photosystem II and also gains protons (H+) from the stroma
2. When PQ donates electrons to the cytochrome complex, it also releases protons into the lumen, increasing the proton concentration there
The chloroplast's ATP synthase is identical to the ATP synthase used in cellular respiration
Cyclic Electron Flow
1. Photosystem I can function independently of photosystem II
2. Reduced ferredoxin donates electrons back to plastoquinone
3. Plastoquinone gets continually reduced and oxidized, keeping protons moving across the thylakoid membrane without the involvement of photosystem II
Electron transfer from the cytochrome complex and shuttling by plastocyanin
Electrons pass from the cytochrome complex to the mobile carrier plastocyanin, which shuttles electrons to photosystem I
The net result of cyclic electron transport is that the energy absorbed from light is converted into the chemical energy of ATP without the oxidation of water or the reduction of NADP+ to NADPH