In fishes, each rib has a single head that articulates with the vertebra at the basapophysis. In tetrapods, the ventral ribs were lost and the dorsal limbs evolved into trunk ribs. Trunk ribs have two heads that articulate with the vertebrae. The tuberculum is the dorsal head that articulates with the diapophysis, and the capitulum is the ventral head that articulates with the parapophysis.