Key derived traits of Angiosperm: "three Fs"—flowers, double fertilization, and fruits
Flower Structure and Function
Floral organs—sepals, petals, stamens, and carpels—are attached to a part of the stem called the receptacle
STAMEN: Filament & Anther – microsporangia (pollen sac that produce pollen)
CARPEL: Style, Stigma & Ovary (ovules)
PISTIL: the term sometimes used to refer to a single carpel or two or more fused carpels
Flower
Complete flowers have all four basic floral organs
Incompleteflowers - lacking sepals, petals, stamens, or carpels
Inflorescences – showy clusters of flowers
Development of Male Gametophytes in Pollen Grains
1. Development of Female Gametophytes (Embryo Sacs)
2. Pollination: the transfer of pollen from an anther to a stigma
3. Coevolution of Flower and Pollinator
Double Fertilization
Seed Development, Form, and Function
Endosperm Development
Endosperm usually develops before the embryo does
Embryo Development
Terminal cell gives rise to most of the embryo
Structure of the Mature Seed
The embryo, which is surrounded by a food supply (cotyledons, endosperm, or both), enters dormancy
Seed Dormancy
An Adaptation for Tough Times: Environmental conditions required to break seed dormancy vary among species
Seed Germination and Seedling Development
Imbibition - the uptake of water due to the low water potential of the dry seed
Fruit Form and Function
Fruit - protects the enclosed seeds and, when mature, aids in their dispersal by wind or animals
Asexual Reproduction
When offspring are derived from a single parent without fusion of egg and sperm
Mechanisms of Asexual Reproduction
Fragmentation - the separation of a parent plant into parts that develop into whole plants
Apomixis - asexual production of seeds
Advantages and Disadvantages of Asexual Versus Sexual Reproduction
There is no need for a pollinator
Allows the plant to pass on all of its genetic legacy intact to its progeny
Can potentially clone many copies of itself
In stable environment: progeny produced by asexual reproduction are stronger than seedlings produced by sexual reproduction
Offspring usually arise from mature vegetative fragments from the parent plant (vegetative reproduction)
Mechanisms That Prevent Self-Fertilization
Dioecious species - plants cannot self-fertilize because different individuals have either staminate flowers (lacking carpels) or carpellate flowers (lacking stamens)
Plants that have flowers with functional stamens and carpels that mature at different times or are structurally arranged in such a way that it is unlikely that an animal pollinator could transfer pollen from anther to a stigma of the same flower
Self-incompatibility - the ability of a plant to reject its own pollen and sometimes the pollen of closely related individuals
S-genes
Vegetative Propagation and Agriculture
Clones from Cuttings
Grafting (Stock & Scion)
Test-Tube Cloning and Related Techniques – in vitro methods
Sexual Reproduction
The fusion of haploid gametes forms a diploid cell, the zygote
Asexual Reproduction
The generation of new individuals without the fusion of egg and sperm
Mechanisms of Asexual Reproduction
Fission - the separation of a parent organism into two individuals of approximately equal size
Budding - new individuals arise from outgrowths of existing ones
Fragmentation -- the breaking of the body into several pieces, followed by regeneration -- the regrowth of lost body parts
Parthenogenesis is asexual reproduction in which an egg develops without being fertilized
Sexual Reproduction: An Evolutionary Enigma
Reproductive Cycles: Ovulation - the release of mature eggs, occurs at the midpoint of each cycle. Reproductive cycles are controlled by hormones, which in turn are regulated by environmental cues.
Variation in Patterns of Sexual Reproduction
Hermaphroditism -- in which each individual has both male and female reproductive systems
Sex reversal – change in sex
Fertilization
External fertilization - the female releases eggs into the environment, where the male then fertilizes them. Internal fertilization: Sperm are deposited in or near the female reproductive tract, and fertilization occurs within the tract.
Ensuring the Survival of Offspring
Internal fertilization is typically associated with the production of fewer gametes but the survival of a higher fraction of zygotes. Internal fertilization is also more often associated with mechanisms that provide greater protection of the embryos and parental care of the young.
Courtship behavior
It allows mate choice and, by triggering the release of both sperm and eggs, increases the probability of successful fertilization.
Pheromones
Chemicals released by one organism that can influence the physiology and behavior of other individuals of the same species.
Gamete Production and Delivery
Sexual reproduction in animals relies on sets of cells that are precursors for eggs and sperm. Gonads - the organs that produce gametes in most animals. Sets of accessory tubes and glands that carry, nourish, and protect the gametes and sometimes the developing embryos. Spermathecae - sacs in which sperm may be stored for extended periods, a year or more in some species.
Cloaca
Common opening of digestive, excretory, and reproductive system.
Monogamy, the sustained sexual partnership of two individuals, is relatively rare among animals, including most mammals.
Ovaries, Oviducts and Uterus, Vagina, Mammary Glands
Male Reproductive Anatomy
External and Internal reproductive structures
External Reproductive Structure
Penis
Internal Reproductive Structure
Testes, Ducts, Accessory Glands
Gametogenesis
Spermatogenesis, the formation and development of sperm, is continuous and prolific in adult males. Oogenesis, the development of mature oocytes (eggs), is a prolonged process in the human female.
Regulation of Mammalian Reproduction
Interplay of tropic and sex hormones
Hormonal Control of Female Reproductive Cycles
Menstrual cycle or uterine cycle (average 28 days), Ovarian cycle