Have significant and disproportionate effect on the community
Not only affects species that it has a direct relationship with, but also those other species that it no longer has direct contact with
Very strong cascading impacts on the structure and functioning of ecosystems
Its loss may lead to the collapse of the community
Keystone species
Old World fruit bats as pollinators
Sea otter (Enhydra lutris) as a keystone predator feeding on herbivorous sea urchins
Ecological niche
Biological role played by a species in the environment
Place or function of a given organism within its ecosystem
If no competition, organism occupies fundamental niche
With competition, organism occupies realized niche = conditions under which an organism actually lives
Ecological Succession
Gradual changes of the vegetational community over time
Characterized by the replacement of early seral stage pioneer or opportunistic species by late-stage equilibrium species
In response to changes brought about by vegetation itself
Stages of succession
Nudation - colonization of a disturbed area
Invasion - arrival of propagules/seeds of various organisms and their establishment in the new area
Competition and Reaction - replacement of more competitive, longer-lived species such as shrubs, and then trees
Stabilization and Climax - further changes take place very slowly, site is dominated by long-lived, highly competitive species
Forms of disturbance
Large-scale disturbance - severe disturbance over a large area results in the colonization of the site by opportunistic species
Small-scale disturbance - small gaps typically result in reorganization of vegetation, important in maintaining diversity in tropical rainforest, results in patches of different stages of successional or compositional maturity, promotes species richness and structural diversity
Climax stage of succession
An equilibrium or steady state with the environment
Last stage is mature, self-sustaining, self-reproducing through developmental stages, and relatively permanent
Vegetation is tolerant to changes in environmental conditions
Theoretical approaches to climax
Monoclimax Theory - only one climax, determined solely by climate, with uniform plant and animal composition
Polyclimax Theory - mosaic of vegetational climaxes controlled by soil moisture, soil nutrients, topography, slope exposure, fire, and animal activity
Climax Pattern Hypothesis - composition, species structure and balance of a climax community are determined by the total environment of the ecosystem