1. No soil, extremes of temperature and water availability, first colonisers are simple autotrophs like algae and lichens
2. Mosses colonise, thin layer of soil builds up
3. Grasses and ferns colonise
4. Soil builds up, abiotic factors become less extreme, seedlings of less hardy plants can survive
5. Flowering plants colonise once insects that pollinate them are established
6. Trees colonise when soil is deep enough and edaphic factors are suitable