chemicals released by one individual and which can affect the behaviour or physiology of another
tropism
directional growth response in which the direction of the response is determined by the direction of the external stimulus
phototropism
shoots grow towards light (they are positively phototropic) which enables them to photosynthesise
geotropism
roots grow towards the pull of gravity
this anchors them in the soil and helps them take up water which is needed for support (keeps them turgid) as a raw material for the plant and to help cool the plant.
minerals such as nitrate in water needed for synthesis of amino acids
chemotropism
pollen tubes grow down the style, attracted by chemicals, towards the ovary where fertilisation can take place
thigmotropism
shoots of climbing plants wind around other plants or solid structures to gain support
nastic responses
non-directional responses to external stimuli
if plant responds towards a stimulus - positive tropic response (directional)
if plant responds away from a stimulus - negative tropic response (directional)
cytokinins
promote cell division
delay leaf senescene
overcome apical dominance
promote cell expansion
abscisic acid
inhibits seed germination and growth
causes stomatal closure when the plant is stressed by low water availibility
auxins
promote cell elongation
inhibit growth of side shoots
inhibit leaf abscission (leaf fall)
gibberellins
promote seed germination and growth of stems
Ethene
promotes fruit ripening
How plant hormones work
produced in a variety of tissues in the plant
hormones reach their target cells
they bind to receptors on plasma membrane
specific hormones have specific shapes which only bind to specific receptors with complementary shapes on membranes of particular cells
hormones can influence cell division, cell elongation and cell differentiation
role of auxins
plant hormones responsible for growth
growth of plant upwards leads from the top apical stem
all buds below apical bud (lateral buds) are inhibited - known as apical dominance
more auxin - slows lateral growth
less auxin - speeds up lateral growth
when apical tip is removed auxin levels drop. lateral growth then occurs - cut end might have produced a hormone that promoted lateral growth
with auxin transport inhibitor placing a ring of auxin transport inhibitor that prevents auxin growth on the stem also results in lateral growth
auxin paste applied to a cut shoot, lateral buds did not grow
scientist the applied an auxin transport inhibitor below apex of shoot and lateral buds grew
abscisic acid
lateral bud growth inhibition - slows growth outwards. high auxin levels encourage high abscistic acid levels
cytokinins
bud growth promotion - travel to where auxin is more concentrated
role of auxins, cytokinins and abscisic acid to apical dominance ( with plant tip intact)
high auxin levels
high abscisic acid levels
low cytokinins in lateral buds
inhibit lateral bud growth
role of auxins, cytokinins and abscisic acid to apical dominance ( with plant tip removed)
low auxin levels
low abscisic acid levels
higher cytokinin levels in lateral buds
lateral buds grow
testing gibberellic acid on different plants
fungal disease in Japan called bakanae was causing rice plants to grow very tall.
Gibberellic acid (GA3) was found in the fungus
Gibberellins responsible for stem elongation and seed germination
2. Testing GA1 levels
compared GA1 levels of tall pea plants an dwarf pea plants which were otherwise genetically identicsl
they found that plants with higher GA1 levels were taller
3. showing GA1 directly causes stem elongation
researchers need to know how GA1 is formed.
They worked out that Le gene was responsible for producing the enzyme that converted GA20 to GA1
researchers then chose a pea plant with a mutation that blocks Gibberellin production between ent-kaurene and GA12 - aldehyde in the pathway
those plants produce no gibberellin and grow to only about 1cm tall
investigating the effect of plant hormones on phototropisms
experiments to confirm the role of the shoot tip in producing a chemical messenger that controls phototropic responses
Boysen Jensen's work
confirmed that water and/or solutes need to be able to move backwards from the shoot tip for phototropism to happen
when a permeable gelatine block inserted behind shoot tip, shoot still showed positive phototropism
when impermeable mica block was inserted, there was no phototropic response