A vascular network runs through the mesophyll, providing the cell walls with water and removing the food products of photosynthesis to other parts of the plants.
The primary photosynthetic organ of most plants, leaves are usually flattened blades that consist, internally, mostly of parenchyma tissue called the mesophyll, which is made up of loosely arranged cells with spaces between them.
Vessels usually are shorter and broader than tracheids, and in addition to pits they have perforation through which water and dissolved nutrients may freely pass.
Associated with the sieve elements are companion cells that do contain nuclei and that are responsible for manufacturing and secreting substances into the sieve elements and removing waste products from them.
Two types of sieve elements occur: sieve cells, with narrow pores in rather uniform clusters on the cell walls, and sieve-tube members, with larger pores on some walls of the cell than on others.
Collenchyma tissue is made up of cells with unevenly thickened primary cell walls and is pliable, functioning as support tissue in young, growing portions of plants.
These organs all contain the three kinds of tissue systems mentioned above, but they differ in the way the cells are specialized to carry out different functions.
The principal cells of phloem, the sieve elements, are so called because of the clusters of pores in their walls through which the protoplasts of adjoining cells are connected.
The vascular tissue system consists of two kinds of conducting tissues: the xylem, responsible for conduction of water and dissolved mineral nutrients, and the phloem, responsible for conduction of food.