A typical eukaryotic cell consists of a cell surface membrane that defines the boundary of a cell and retains its contents, a nucleus that contains the genetic material that directs cellular activities, and cytoplasm that consists of cytosol, the aqueous solution of ions and organic compounds (e.g., sugars, amino acids, and proteins), and organelles, which include the organic cellular structures, usually enveloped by a membrane, such as the ribosome, endoplasmic reticulum, Golgi body, lysosome, mitochondrion, vacuole, and centriole.