-begins with synapsis-DNA molecule of one of the chromatids is cut and a second cut is made at the same point in the DNA of a non-sister chromosome -the DNA of each chromatid is joined up to the DNA of the non-sister chromosome, swapping sections of DNA between chromatids-later in prophase I, the tight pairing of the homologous chromosomes ends, but sister chromatids remain tightly connected→chiasma-results in recombination