Two-legged (bipedal) vertebrates Distinguished from other vertebrates by: • Feathers • filamentous, soft, flexible, light-weight structures • dead when functional – must be replaced regularly•
Skull articulates with the first neck vertebra by means of a singleoccipitalcondyle
Simple middle ear with only 1bone (stapes) mammals have 3
birds and reptiles have nucleated red blood cells
cursorial theory - running dinosaurs heldout arms to heighten leaping ability and catch insects • Could jump higher and help maintain balance
arboreal theory - climbing dinosaurs heldout arms to control leapsfrom branch to branch• Gliding/parachuting
• 2 hypothesis about how they evolved:• Feathers aided in flight – elongated, frayedscales may have enhanced gliding/parachuting • temperature regulation devices • Some say it acted as heatshield that allowed birds to be more active in hot environments • Others say – prevent heat loss (endothermic dinosaur ancestors)
feathers are composed of beta-keratin (have some similarities to alpha-keratin which is found in skin of all vertebrates – but not in the same group of proteins)
The beta-keratin found in feathers belongs to a special class of proteins characterized by a deletion in their molecular sequence
shaft down the center with the flexible vane on either side
• The shaft has two parts: • calamus (quill) – short, tubular base which stays attached to the follicle until it is molted • rachis – long, central shaft which supports the vein
The feather vane is made up of hundreds of tiny parallel barbs branch off the rachis
barbules branch off the barbs which have hooklets that hold the feather together
• Further out, the vane is firm, compact, and closelyknit (pennaceous)• Used as an airfoil• Protects the undercoat• Sheds water• reflects/absorbs solar radiation• Used in visual and auditory communication
At the base of the vane are soft, loose, fluffy barbs(plumaceous) used for insulation
• Contour feathers – include typical body feathersand flight feathers• Wing feathers – remiges (remex)• Tail feathers – rectrices (rectrix)