4.3 - 3.8 Ga was a period of intense comet and asteroid bombardment
Precambrian Life
Earliest fossil is 3.55 Ga but chemical signatures of biological origin have been dated to older rocks
Stromatolites 3.8 Ga - formed by a growth of layer upon layer of photosynthetic bacteria
Banded Iron Formations appear at around 3.7 Ga and they indicate that the great oxygenation event was at 2.4 Ga
Eukaryotes appear around 1.8Ga
Ediacaran fauna appear around 565 Ma
Cambrian Explosion
Explosion of life in the early Cambrian
Most major phyla appear during this time
Major diversification of life
Hard parts appear
Factors causing the Cambrian Explosion
Oxygen levels - increase in oxygen levels lead to an increase in size, ozone filtered out harmful UV rays allowing complex life
CaCO3 Concentration - increase in CaCO3 provided material for hard parts
Predation - created selection pressures and also caused eyes and burrowing behaviour
Rise in sea level provided more ecospace
Hard parts - more efficient predation, protection for prey, efficient movement and stronger body for protection against currents
Preservation bias - appearance of hard parts increases chance of preservation - may not be real
Evidence for the Cambrian Explosion
Almost every phylum known appears in the Cambrian
Trace fossils - show an increased diversity in Cambrian rocks and development of new ecological niches and strategies
Why did hard parts evolve?
Protection from predators
Mineralised mouth parts for eating
To facilitate movement and exploration of new environments
Protection from the environment
Support structures for soft tissue allowing larger sizes
Ordovician Radiation
Ecosystems increased in complexity
Fauna emerging in the Ordovician set the template for the rest of the Palaeozoic
Ordovician adaptive radiation - increase in diversity, particularly in filter feeding organisms
Explain what is meant by the term Cambrian Explosion
The sudden appearance of hard parts and organisms such as trilobites and brachiopods. It was an increase in diversity of species and fossils.
A increase in chemical weathering releases calcium carbonate for use in hard parts of organisms. Chemical weathering adds more nutrients into oceans allowing more metabolic processes/building bigger bodies
Glaciations/meteorite impacts lead to mass extinctions
Rapid and dramatic environmental catastrophes allow for subsequent rapid diversification of life due to innovation and development of new animal groups and competition and selective processes following obliteration of previous ecosystems
Suggest why it is difficult to reconstruct ecosystems occurring in the Precambrian
It was a long time ago - greater chance for metamorphism, weathering or erosion which may have destroyed fossils. There were no hard parts then, meaning preservation potential was low. Furthermore, there are no living forms present today meaning we cannot apply uniformitarianism, and the diversity of life was low meaning the chance of being preserved was lower
The Rise of Tetrapods
Tetrapod - four limbed animal containing extinct and extant amphibians, reptiles, synapsids (incl. mammals) etc
Amphibians - tetrapods with lifestyles in and out of water. Evolved from lobe-finned fish in the late devonian to early carboniferous.
Lobe finned fish - four fleshy fins supported by a bone in a similar structure to a hand
Tiktaalik - fossil intermediate between fish and tetrapods - specimens are extremely well preserved allowing detailed comparisons between lobe finned fish and early amphibians
Lobe finned fish to early amphibians - differences
Development of skeletal girdle connecting limb bones to the skeleton for better movement on land
More robust skeleton strengthening vertebral column and rib bones - extra support on land
Eyelids formed to keep eyes moist as no longer submerged in water
Development of a double circulatory system
Tongue formed within its mouth to catch prey
Ears to detect sound waves through the air
Slender skull
Evolution on land
Amniotic egg - provided protection whilst remaining porous allowing diffusion of gases into and out of egg (respiration)
Yolk sac and albumin - Yolk sac - food, albumin - water nutrients
Amnion - fluid filled sac around embryo
Amniotic egg allowed amniotes to move away from bodies of water and become larger
Rise of dinosaurs and mammals
Dinosaurs evolved from archosaurs - after permio-triassic mass extinction
Birds are throught to have evolved from theropod dinosaurs in the late Jurassic. Similarities between dinosaurs and birds include:
Hollow thin-walled bones
S shaped curved neck
Elongated arms and forelimbs
Pubis shifted from anterior (forward position) to a posterior (backward position)
Large eye sockets in the skull
Hinged ankles to reduce rotation
Archaeopteryx
Semi arboreal animal
Capable of gliding and sustaining flight
Shows characteristics of dinosaurs AND birds
Feathers
Earliest preserved feathers are found in the late Jurassic
Feathers evolved to be more complex, becoming elongate and symmetrical
Mammals
225 Ma mammals and dinosaurs appear in the rock record
Many mammals lived in burrows and were nocturnal
During the mesozoic mammals yielded a vast array of body forms
Land mammals kept getting larger for 35Ma after the dinosaurs were wiped off the planet
Homosapiens appear around 300Ka
Characteristics of a Mass Extinction
Large drop in diversity
High number of extinctions at the same time
Extinction rate is higher than origination rate
Wide variety of groups affected
Geographically widespread
Short time span
Sharp recovery - new fauna fills available ecospace
Fern spike common on land
How can extinctions be selective?
Size
Abundance
Distribution
Specialisation
Adaptability
Morphology and mode of life
Cretaceous - Palaeogene
17% of families, 50% genera, 75% of all species
Non-avian dinosaurs and ammonites extinct
Echinoids, bivalves and fish severely affected
Mammals, birds, crocodiles survive
Mammals and birds radiate
Evidence for the K-Pg Mass Extinction caused by meteorite
Iridium - layer concentrated at K-T boundary and most iridium comes from space
Shocked quartz forms under intense pressure and is associated with meteorite craters
Tektites - gravel sized glass formed from melting of soil and rock
Sedimentary evidence suggests a large Tsunami reached Texas
Chicxulub impact crater dated 65Ma
Deccan traps - widespread flood basalts in India. Millions of tons of sulphur dioxide
Consequences of meteorite impact (K-Pg extinction)
Would have thrown up dust into the atmosphere as well as releasing sulphur dioxide
Resulting in widespread cooling + acid rain
Resulting in the collapse of life and the rest of the food chain
Fern spike due to being able to tolerate acidic conditions.
Permian - Triassic
57% families, 83% genera, 95% marine species
Trilobites, rugose and tabulate corals extinct
Most brachiopods extinct
Ammonoids and echinoids severely affected
Land reptiles and many trees extinct
Evidence for the P-T mass extinction
Sedimentary evidence - widespread deposition of black shales and iron pyrite suggesting marine anoxia
Tectonic - Pangaea provided opportunity for glaciers to form.
Shocked quartz - found in Australia and Antarctica. However recently refuted suggesting that it is due to plastic deformation associated with tectonic activity
Meteorite impact crater - Several impact sites been suggested but later refuted.
Siberian traps - largest flood basalt provinces on Earth - time interval of around 900K years. Worldwide atmospheric contamination.
Macroevolution and the tree of life
Life on earth is extremely varied - genetic and morphological data suggests all life on Earth is related
Tree of life shows phylogeny of organisms
Large trends and transformations in evolution above the species level is termed macroevolution
Evolution is: descent with modification OR genetics + time
Phyletic Gradualism
Traditional view of evolution
Gradual change
Slow, steady divergence of lineages
Many transition forms are not preserved which makes evolution appear faster
Punctuated equlibrium
Long periods of stasis/stability
A large amount of change in a short time, tied to a speciation event
Transitional forms exist for a short period
Evidence for punctuated equlibrium - variation in the number of eye lenses of the trilobite genus
More evidence for punctuated equilibrium includes:
Cambrian explosion/radiation
Radiation of mammals
Living fossils such as nautilus
Cladistics
Aims to show evolutionary relationships based on physical traits shared by different groups of organisms
Climate
Sum of weather conditions in a region over a long time, including variation in temperature, humidity, atmospheric pressure, wind and precipitation
Tilt of the Earth results in seasonality
Gulf stream - long stream of warm surface water that keeps Europe warm and north America cold. Partly caused by circulation of ocean waters but mainly driven by winds
Global circulation model - cooling effects around the globe driven by wind. Main source of heat is energy from short wave light from the sun. Heat comes from light absorbed from Earth’s surface re-emitted as long wave radiation, heating the air around it. Excess of heat at the equator and deficit at the poles due to the amount of sunlight hitting them (more at equator, less at poles).
Ocean current circulation - driven by wind, tides, changes in water density and rotation of the earth. Modified by topography of ocean floor. Surface currents control motion of top 10% of ocean’s water. Deep ocean currents control other 90%.
What causes climate change?
Changes in atmospheric composition of greenhouse gases
greenhouse effect - Atmosphere traps sun’s energy - EM radiation in broad range of wavelengths. Rock, soil and water reemit energy as heat.
Carbon dioxide, methane, water vapour, nitrous oxide - absorb IR radiation and re-emit. Not much escaping radiation leaves without encountering greenhouse gases. Greenhouse gases maintain the average surface temperature by retaining heat energy.
Volcanic activity, methane hydrates and chemical weathering all influence atmospheric greenhouse gas composition
What causes climate change?
Distribution of the continents
Determines route of global current systems and strength of oceanic heat exchange between poles and equator. Movement of continents determines long term global climate
The carboniferous period - near continuous continental landmass extending from North to South poles.
Late carboniferous icehouse - development of ice sheets occurred due to:
Circulation of polar and tropical waters restricted by landmasses