Thin-walled region between epidermis and endodermis
Contains parenchyma, collenchyma and sclerenchyma tissues
Distinct layer in dicots, not in monocots
Contains starch grains, oil, tannins, crystals
May develop chloroplasts and become photosynthetic (chlorenchyma)
In hydrophytes, may be aerenchymatous (spongy with air spaces)
Contains special cells like sclereids, resin ducts, oil glands, laticifers