Anthers and ovules bear sporangia, structures where spores are produced by meiosis and gametophytes develop. Pollen grains, each consisting of a mature male gametophyte surrounded by a spore wall, are formed within pollen sacs (microsporangia) of anthers. An egg-producing female gametophyte, or embryo sac, forms within each ovule. In angiosperms, pollination is the transfer of pollen from an anther to a stigma. If pollination is successful, a pollen grain produces a structure called a pollen tube, which grows and digests its way down into the ovary via the style and discharges sperm in the vicinity of the embryo sac, resulting in fertilization of the egg.