Cnidaria are diploblastic animals; in other words, they have two main cell layers, while more complex animals are triploblasts having three main layers
The two main cell layers of cnidarians form epithelia that are mostly one cell thick, and are attached to a fibrous basement membrane, which they secrete
They also secrete the jelly-like mesoglea that separates the layers
Epitheliomuscular cells whose bodies form part of the epithelium but whose bases extend to form muscle fibers in parallel rows
The fibers of the outward-facing cell layer generally run at right angles to the fibers of the inward-facing one
In Anthozoa (anemones, corals, etc.) and Scyphozoa (jellyfish), the mesoglea also contains some muscle cells
Cnidocytes, the harpoon-like "nettle cells" that give the phylum Cnidaria its name, appear between or sometimes on top of the muscle cells
Nerve cells. Sensory cells appear between or sometimes on top of the muscle cells, and communicate via synapses (gaps across which chemical signals flow) with motor nerve cells, which lie mostly between the bases of the muscle cells
Interstitial cells, which are unspecialized and can replace lost or damaged cells by transforming into the appropriate types, are found between the bases of muscle cells