Have a "true" body cavity that is completely surrounded by mesoderm
Coelom
Closed, fluid filled cavity that surrounds the gut
The fluid within acts as a circulatory system (blood)
Tissue membranes (mesenteries) suspend organs in the coelom
Metamerism
The body is made up of a series of repeating, coordinated segments called metameres that are separated from one another by septa
Each metamere contains sets of repeating organs
Annelid body
Two part head consisting of Prostomium and Peristomium
Annuli: circular rings
Pygidium: the last segment; where the anus is
Clitellum: reproductive structure
Annelid nervous system
2 cerebral ganglia
A ventral nerve cord with 2 ganglia per metamere
Some species have sensory organs such as eyes, palps, and tentacles
Annelid locomotion
Both longitudinal and circular muscles which allow them to "wiggle"
Most have setae (chitinous bristles secreted by the epidermis) that aid in locomotion and burrowing
Fluid in coelom acts as a hydrostatic skeleton
Annelid gas exchange
Mainly by diffusion through skin
Class Polychaeta often has specialized structures for gas exchange (e.g. parapodia, gills)
Annelid digestive system
Unidirectional
Regional specialization (digestive organs)
Annelid circulatory system
Closed circulatory system composed of blood vessels (some of which are contractile and act as "hearts")
Some circulation is also accomplished by the coelomic fluid
Annelid excretion
Excretion is accomplished by organs called nephridia (singular nephridium)
Annelid reproduction
Asexual (budding or fission), and sexual
Class Polychaeta
All marine
Contains 2/3 of all known Annelids
Common species are clamworms, sandworms, and lugworms
Have a well developed head with specialized sense organs
Polychaeta
Commonly called "bristle worms"
Have many setae (chitinous bristles secreted by the epidermis) arranged in bundles on paddle-like appendages called parapodia
Parapodium
Function in gas exchange, locomotion, and feeding
Tagmatization (tagmosis)
The fusion and specialization of formerly metameric segments
Polychaeta
Many are filter-feeders with specialized structures
Many are predatory with specialized structures
Many construct their own homes out of CaCO3 or sand debris and mucous
Polychaeta reproduction
Usually dioecious (separate male and female)
No permanent sex organs; gametes are shed into coelom where they stay until time for fertilization
Fertilization is usually external in water
Indirect development - trocophore larvae swims around until it grows into adult form by adding segments
Polychaeta defense
Some have tubes to hide in
Some have vicious jaws
Some have modified "stinging" setae
Some Polychaetes have a mutualistic relationship with their host, such as scale worms found near, or in the mouth, of brittle stars, starfish, and sea urchins
Class Oligochaeta
Habitats: terrestrial, freshwater, and marine
Have few setae (Oligo = few, chaeta = setae)
What do earthworms eat?
Usually feed on detritus (decaying organic matter)
Can feed on dead leaves and plant roots, living things such as other worms, bacteria, and fungi, and decomposing remains of other animals
Earthworms consume dirt as they burrow, but they do not get nutrients from the dirt - their nutrients come from decaying plant and animal remains that are in the dirt
Earthworm digestive system
Specialized to obtain the maximum amount of nutrients out of the detritus, e.g. pharynx, crop, gizzard
Earthworm locomotion
Circular muscle contraction and longitudinal muscle contraction
Earthworm reproduction
Usually monoecious, still have to have a partner to cross-fertilize by exchanging sperm
Earthworms are essential soil aerators, mixing the soil with their tunnels and providing fertilizer through their feces
Class Hirudinea
Includes leeches
Usually freshwater but there are some marine and terrestrial species
No septa between metameres
No setae or parapodia
Have 2 suckers
Hirudinea
Have an extendable proboscis for feeding
Usually have a fixed number of segments (34)
Each metamere consists of several annuli
Hirudinea locomotion
Lack septa between metameres, so they are incapable of moving like Oligochaetes - instead, they use their anterior and posterior suckers to move
Hirudinea reproduction
Usually monoecious, cross-fertilize by exchanging sperm
Although some leeches are parasitic blood suckers, many are predators
Leeches have been used medicinally since the 19th century, currently used to increase blood flow following reconstructive surgery, and their salivary glands contain the powerful anticoagulant hirudin