A collection of species or populations, and interactions between these collections over time, and in a given space
No two communities will be exactly alike even if we have the same number of organisms of each species of the same size and age in the same position etc.
The challenge is to be able to generalise to the point can make useful comparisons / understand systems, without being so sweeping as to eliminate the subtlety
Communities are more of a continuum than a discreet entity
Communities as groups of species
A useful definition only in that it is relatively simply to define, but it ignores the components of space
Communities as a collection of populations
Plant community data typically looks at numbers of individuals within species whereas animal data focuses on the presence or absence of species
Abstract vs concrete community
The concrete form refers to it at a given point, or components of it that exist. The abstract community is the more generalised version
Ways communities may show organisation (structure)
Species composition
Temporal
Trophic
Species composition
At any given time a community will have a pool of species represented, with dominant and keystone species
Keystone species
Those on which others (or even the community as a whole) depends, and their loss will lead to the loss of others
Indicator species
Those which provide information about a community and can be used as a proxy or marker for the likely condition of the community
Temporal structures
Diurnal
Nocturnal
Crepuscular
Seasonal
Communities change over time (succession) and can be long term (millions of years) or daily (intertidal zones) and individuals may live for days or centuries
Different species occupy landscapes at very different scales though may still interact (krill vs whales)
Predation
The complete consumption of one organism by another, where one is harmed and one benefits
Parasitism
Consumption of part or all of an organism by another, where one is harmed and one benefits, usually lives on / in the host
Mutualism
An interaction where both organisms benefit
Food web
Illustrates relationships (what eats what) between species or populations within a community, and represents energy transfer
Energy enters ecosystems as light, and leaves primarily as heat
Trophic levels
Primary producers
Primary consumers
Secondary consumers
Definitions of trophic levels are not as clear cut as we would like, as some organisms can be in multiple levels
On average, only 10% of the energy makes it from one trophic level to the next
Ways to represent transfer of energy or trophic levels
Pyramids of numbers
Pyramids of biomass
Food chains or webs
Food chains are typically short (few have more than 3 or 4 links) or the system would run out of energy to support higher trophic levels
Selection may potentially increase and decrease food chain lengths
Guilds
Aggregations of single species or populations into similar functional types for simplicity
Compartmentalisation
Webs can be compartmentalised into groups where interactions are strong, but between which interactions are weak
Ways to trace food webs
Observational data of feeding
Examination / analysis of waste
Isotope tracking
Ecotron
Food webs can be used to help build models of interactions and predict effects of changes, which is increasingly important for conservation and with problems like invasive species