Supportive system that provides protection, support, and a place for muscle attachment
Hydrostatic skeleton
Muscles in the body wall develop force by contracting against incompressiblecoelomic fluids
Alternate contractions of circular and longitudinal muscles of the body wall enable a worm to move forward
Muscular hydrostats
Composed of incompressible tissues
Complex movements are a result of complex arrangements of muscles
Muscular hydrostats
Elephant's trunk
Mammal & reptile tongues
Cephalopod tentacles
Rigid skeletons
Contain some kind of rigid elements
Types of rigid skeletons
Exoskeleton
Endoskeleton
Exoskeleton
Mostly made of chitin, found in molluscs & arthropods and some other invertebrates
Endoskeleton
Found in echinoderms, chordates, and some cnidarians, composed of two types of tissues (bone and cartilage)
Vertebrate endoskeleton
Composed of bone and cartilage (types of connective tissue)
Bone provides support, protection, and serves as a reservoir for calcium and phosphorous
Notochord
Supportive rod found in protochordates and developing vertebrates, derived from mesoderm, except in jawless vertebrates it is replaced by the backbone
Jawless fishes and elasmobranchs have cartilaginous skeletons - a derived feature since their ancestors had bony skeletons
Most vertebrates have bony skeletons, with some cartilaginous parts
Cartilage
Soft, pliable tissue that resists compression and is variable in form
Hyaline cartilage has a clear, glassy appearance with chondrocytes surrounded by a matrix, no blood vessels
Cartilage is often found at articulating surfaces of many bone joints, and as supporting rings of the passageways in the respiratory system
Cartilage similar to hyaline cartilage is found in many invertebrates, such as the radula of gastropods and the lophophore of brachiopods
Bone
Highly vascular living tissue that contains significant deposits of inorganic calcium salts
Types of bone
Endochondral (replacement) bone
Intramembranous bone
Endochondral (replacement) bone
Develops from another form of connective tissue - usually cartilage
Intramembranous bone
Develops directly from sheets of embryonic cells, found in the face, cranium, clavicle, and dermal bone
Types of bone density
Spongy bone
Compact bone
Spongy bone
Consists of open, interlacing framework of bony tissue, oriented to give strength
Compact bone
Dense, the open framework of spongy bone has been filled in by additional calcium salts, composed of a calcified bone matrix arranged in sets of concentric rings - osteons
Bone
Bundles of osteons interconnected with blood vessels and nerves
Between the rings are lacunae (cavities) filled with osteocytes (bone cells) connected by tiny passageways that distribute nutrients
Bone remodeling
Osteoclasts are bone resorbing cells, osteoblasts are bone building cells, both processes occur together so that new osteons are formed as old ones are resorbed
Hormones (parathyroid hormone for resorption and calcitonin for deposition) are responsible for maintaining a constant calcium level in the blood
Vertebrate skeleton
Axial skeleton
Appendicular skeleton
Axial skeleton
Includes the skull, vertebral column, ribs, and sternum
Appendicular skeleton
Includes the limbs and pectoral and pelvic girdles
Over time, the number of skull bones has been reduced from as many as 180 in some early fishes to 35 or fewer in mammals
Vertebral column
Serves as the main stiffening axis, provides points for muscle attachment, provides stiffness, and preserves body shape during muscle contraction much like the notochord from which it is derived
Most vertebrates have paired appendages
Pectoral and pelvic fins
In fishes, supported by the pectoral and pelvic girdles
Tetrapods
Have two pairs of pentadactyl limbs (although they may be highly modified through bone loss or fusion)
Pelvic girdle
Generally firmly attached to the axial skeleton
Pectoral girdle
More loosely attached to the axial skeleton
Animal movement
Most depends on contractile proteins which can change their shape to relax or contract, powered byATP
Cilia
Found throughout the animal kingdom (except in nematodes, rare in arthropods), uniform in diameter (.2-.5 μm) and structure, basal body similar to a centriole - 9 triplets of microtubules composed of the protein tubulin, cilium has 9 pairs surrounding two individual microtubules
Flagellum
A whiplike structure longer than a cilium and usually present singly, structure is the same as cilia but with a different beating pattern