Ventilation is the process of moving air or water in and out of the body
External Respiration - exchange of gases between the atmosphere and the blood
Internal Respiration - exchange of gases between the blood and tissue cells
Respiratory Organs
Cutaneous Respiration (Skin)
Gills
Swim Bladder
Lungs
CUTANEOUS RESPIRATION
The skin is an important supplementary organ for cutaneous respiration in certain organisms. It is typically thin and moist to allow the direct exchange of gases.
The gills are respiratory organs that allow the withdrawal of oxygen in water.
Internal gills: associated with the pharyngeal slits and pouches
External gills: arise from the branchial region as filamentous capillary beds
Swim bladder is an elongated gas-filled sac that aids in respiration. It used to function primarily as a buoyancy organ
Lungs are the primary respiratory organs in terrestrial vertebrates. It enable the extraction of oxygen from the air and expel carbon dioxide.
The Trachea is also known as the windpipe is formed by 16 to 20 stacked, C-shaped pieces of hyaline cartilage that are connected by dense connective tissue
The larynx is cartilaginous structure inferior to the laryngopharynx that connects the pharynx to the trachea and helps regulate the volume of air that enters and leaves the lungs
Swim Bladder
dorsal outgrowth from the pharynx
situated dorsal to the digestive tract
may be paired or unpaired
returning blood drains to the general systemic circulation before entering the heart
Lungs
ventral outpocketing from the floor of the pharynx
situated ventral to the digestive tract
paired organ
venous returns to the heart separately from the generic system circulation
Ciliary pumps facilitate respiratory exchange by ventilating surfaces.
In early vertebrates, they allowed for the evolution of larger species and protective armor by reducing reliance on cutaneous respiration.
Cilia: hair-like projections that line the respiratory routes
create water currents
facilitate feeding by bringing food particles to mucous traps
Surfactants: secretion that lines the lungs and gas bladders
reduces surface tension at the water-air interface
important where there is partitioning of the interior respiratory surface
Muscular pumps facilitate ventilation by contraction and relaxation of respiratory muscles.
Ram ventilation is a technique by which the fish's own forward locomotion contributes to gill ventilation.
Water Ventilation: Dual Pump
Water-breathing fishes utilize a dual pump system comprising buccal and opercular pumps, working synchronously to create a continuous, unidirectional flow of water across the gills
Air Ventilation: Buccal Pump
Air-breathing fishes and amphibians use a buccal pump for lung ventilation, expanding the mouth cavity to draw in air and compressing it to enter the lungs.
Two-stroke: brings air into the mouth → forces it into the lungs in the second
Four-stroke: transfers gas from lungs to mouth → expels air → inhalation of fresh air into the mouth → compression of fresh air into the lungs
Air Ventilation: Aspiration Pump
Air-breathing amniotes employ aspiration pumps to facilitate air entry or exit through suction, utilizing the rib cage and diaphragm
Comparative Anatomy - Osteichthyes
Usually have 5 gill slits
Operculum projects backward over gill chambers
Interbranchial septa are very short or absent
Comparative Anatomy - Agnathans
6 - 15 pairs of gill pouches
pouches connected to pharynx by afferent branchial (or gill) ducts & to exterior by efferent branchial (or gill) ducts
Comparative Anatomy - Amphibians
Lungs of the amphibians have 2 simple sacs; internal lining may be smooth or have simple sacculations
They also relies on cutaneous respiration
Comparative Anatomy - Reptiles
Reptilian lungs are simple sacs in Sphenodon and snakes
Lizards, crocodilians, & turtles - lining is septate, with lots of chambers & sub chambers
air exchanged via positive-pressure ventilation
Comparative Anatomy - Birds
Avian lungs are modified form of reptilian lungs
air sacs (diverticula of lungs) extensively distributed throughout most of the body
arrangement of air ducts in lungs have no passageway or a dead-end
air flow through lungs (parabronchi) is unidirectional
Comparative Anatomy - Mammals
Mammalian lungs are multi chambered & usually divided into lobes
air flow is bidirectional: air exchanged via negative pressure ventilation, with
pressures changing due to contraction & relaxation of diaphragmintercostal muscles
Embryonic Development
The development of the respiratory system begins from the primitive gut tube.
Respiratory diverticulum forms as an outpocketing on the proximal part of the gut tube.