A type of microorganism that lacks a true nucleus and is typically found in soil, water, and the gastrointestinal tracts of animals
The first organisms that appeared on Earth are undoubtedly single-celled, like the prokaryotes
Protists
Exhibits remarkable variety in terms of structure, mode of nutrition, and habitat
Some protists are so simple that they are mistaken as prokaryotes
Others are very huge that they are mistaken as plants
Kingdom Protista is an artificial grouping of organisms coming from different evolutionary lineages
Biologists generally agree that fungi, plants, and animals arose from protist ancestors
Protists
All protists are eukaryotes
They have a well-defined nucleus, and may carry only one nucleus or several nuclei
The protist cell also contains several other membrane-bound organelles, including the endoplasmic reticulum, Golgi complex, lysosomes, mitochondria, and, in some, chloroplasts
They also hold ribosomes and a cytoskeleton made up of microtubules
Unlike plants and animals, the cells of protists are not organized into specialized tissues
Size range of protists
Fractions of a millimeter for a green alga, to about 2 mm diameter for a dinoflagellate, with the notable exception of the giant kelp whose length can reach 65 m
Habitats of protists
Oceans (as part of plankton)
Hydrothermal vents
Rocky intertidal pools
Freshwater ponds
Arable soil
Desert sand
Land covered with ice and snow
Forest litter
Barks of trees
Lifestyles of protists
Heterotrophs
Autotrophs
Symbiosis with other organisms
Parasites
Informal categories of protists
Animal-like protists (protozoans)
Fungus-like protists
Plant-like protists
Animal-like protists (protozoans)
Like animals, protozoans are heterotrophs
They ingest food obtained from the environment which includes bacteria, other protists, and nonliving organic compounds
Unlike animals which are multicellular, protozoans are unicellular organisms
They lack the protection of a cell wall typical of prokaryotes
Habitats of protozoans
Freshwater
Marine waters
Damp soil
Droplets of water on plants (free-living)
Body fluids of living hosts (parasitic)
Parasitic protozoans cause many significant diseases such as malaria and African sleeping sickness
Structures used by protozoans for movement
Flagella
Pseudopodia
Cilia
Some parasitic protozoans lack structures for motility as food is abundant in the surrounding body fluids of the host
Protozoans with flagella (zooflagellates)
Fast swimmers with one or more flagella
The flagella beat rhythmically to push or pull the whole cell through the fluid environment
They generally reproduce by binary fission, producing genetically identical individuals
Protozoans with pseudopodia (amoeba)
Appear as a shapeless mass of cytoplasm bounded by a thin outer cell membrane
The cytoplasmic content streams constantly, presses against the membrane, and creates fingerlike projections called pseudopodia
Pseudopodia aid in the characteristic creeping movement and engulfing of food
Amoeba
The thin outer membrane assists in the exchange of substances between the amoeba and its environment
An organelle called contractile vacuole pumps out excess water and bulky waste materials
Entamoeba histolytica
A parasitic species that causes amoebiasis or amoebic dysentery
Amoebiasis
May be mild or severe
Mild infection includes nausea, loose stool, abdominal tenderness, and occasional fever
Severe infection occurs when the parasite attacks the large intestine and moves beyond, to the liver for example, causing liver abscess
There is a high incidence of amoebiasis in the tropics and subtropics where animal excrement is used as fertilizer for agricultural land
Other protozoans with pseudopodia
Heliozoans
Radiolarians
Foraminiferans
Protozoans with cilia (Paramecium)
The oral groove is lined with cilia that sweep water with suspended food into the mouth pore
Food vacuoles contain the ingested food particles and fuse with lysosomes for digestion
Has a contractile vacuole that prevents the accumulation of excess water inside the cell
Has a large macronucleus that controls many metabolic activities and a micronucleus involved in sexual reproduction
Stentor
A trumpet-shaped freshwater ciliate with cilia arranged in rows lining the entire body and others arranged spirally around its mouth pore, creating feeding currents
Colonial ciliates (Epistylis)
Grows on the outside surface of crustaceans and fishes
Stalked ciliate that can measure from 200 to 250 μm in length, with branched colonies up to 2 mm
Does not harm fishes and crustaceans unless the colonies amass and impede with water flow and draw out dissolved oxygen
Parasitic ciliates (Balantidium coli)
The only ciliate known to parasitize humans
Attacks tissues of the large intestine and causes diarrhea when cysts are ingested from contaminated food and water
Immobile protozoans (apicomplexans)
Lack structures for motility
Have an "apical complex" that enables them to enter a host cell
Are parasites that cause serious human diseases like malaria
Plasmodium
A well-known apicomplexan that causes malaria
Its life cycle involves two different host species - the Anopheles mosquito and humans
Antimalarial drugs are now available but certain Plasmodium strains have become resistant to some of them
Fungus-like protists
Like fungi, they are heterotrophs that decompose dead organic matter
They do not photosynthesize and produce their own food
Their bodies consist of threadlike filaments and they reproduce through spores
Unlike fungi, they produce flagellated cells at some stage in their life cycle
Their cell walls are made up of a different polysaccharide and they contain centrioles
Groups of fungus-like protists
Plasmodial slime molds
Cellular slime molds
Water molds
Downy mildews
Plasmodial slime molds
Colorful fungus-like protists found creeping on decaying logs, moist forest soil and dead leaves
Move by forming pseudopodia and feed on microbes and organic matter
The large, branching, motile structure is known as the plasmodium
When nutrients are depleted, the plasmodium produces stalked fruiting bodies called sporangia with reproductive cells
Cellular slime molds (Dictyostelium discoideum)
Provides a simple model for determining how eukaryotic genes influence many cellular processes
Its complete life cycle, including both unicellular and multicellular stages, can be simulated in the laboratory
Water molds and downy mildews (oomycetes)
Have long, hollow, filamentous bodies containing several nuclei
Their cell wall is composed of cellulose, not chitin as found in true fungi
Produce sporangia that release flagellated reproductive cells called zoospores
Water molds
Aquatic saprophytes that obtain nutrition from decomposing plants and animals in freshwater habitats
Water molds can be seen on dead insects, on salmon after it has completed its spawning run, and on immobile eggs
Dictyostelium discoideum
It is easily grown and its complete life cycle, including both unicellular and multicellular stages, can be simulated in the laboratory