the mouth is opened, increasing volume of buccal cavity
pressure in the cavity drops, water moves into the cavity
opercular valve shuts and opercular cavity containing gill expands, lowering pressure
pressure in buccal cavity increases causing water to move over gills
the mouth closes and opercular opens
pressure in the opercular cavity increases and water is forced over the gills and out the operculum
buccal cavity moved up to maintain a flow of water over the gills