A ranking system where animals are dominant over those below it and submissive to those above it in the heirarchy.
How is a hierarchy maintained?
Ritualised displays and aggression
Ritualised display
Ritualised behaviour by which an animal uses to provide specific information to others
Establishing a hierarchy lowers conflict and aggression in the clan, as the hierarchy is determined early in life during brief fights or threats and displays, after which status is rarely contested except by newcomers.
Advantages of territorialism
Protection from the physical environment
Greate ability to detect predators
Better defence against predators
Greater ability to find resources such as food
Mates available
provision of learning opportunities for young
regulation of population through controlled access to mates/resources
Disadvantages of Territorialism
Increased intraspecific competition
increased risk of contracting and spread of diseases
restricted access to mates
increased risk of infant mortality due to cannibalism
Advantages for omega individuals:
There is a slim prospect of eventual reproduction if one or more of the dominant animals die. For an outcast, the prospects of reproduction may be nil. Lower-order animals also benefit by being members of a group – a group means reduced risk of predation, greater chance of obtaining food
Home range
Geographical area that animals rarely leave, nest in and search within for food and mates
Territory
An area which the animal, breeding pair or group marks and defends
Lek
A territory used only for mating
Polyandry
One female mates with multiple males. Advantage: females can select who to mate with and will often pick a mix of the fittest males. Some females can select which sperm to allow to impregnate her eggs.
Parental care
Any action by which an animal expends energy or assumes risks to benefit its offspring (e.g., nest-building, feeding of young, defence). Increases the chance that offspring will survive but is costly to the parents.
Sexualdimorphism
Differences in physical characteristics between males and females of the same species. Evolves due to intraspecific competition for mates eg. males evolve larger horns to impress females in ritualized fights. Advantage: makes mate selection for the fittest mate easy.
Territorial behaviour
Competitive behaviour in when an animal tries to adopt and defend a physical area against others of the same species.
Omega individual
Most submissive individual in a linear hierarchy
Pair bond
One female mates with one male for their wholelife. Lots of time and energy is invested in courtship and reinforcing the pair bond. Advantage: share energy investment in caring for young, and protecting territory.
Benefits of dominance
-first access to food
-spend less time hunting/foraging
-best access to mates/more mates
-The fittest individuals do the most survival/mating so the best genes spread through the population
Costs of dominance
-Higher metabolic rate so need more food
-have to use energy to defend territories
-high levels of stress hormones
-have to engage in more ritualised fights to defend rank
Dominance
High ranking individuals in a group. Often the strongest/most fit, and have the best access to resources including mates.
Complex hierarchy
A kind of hierarchy where groups of individuals hold the top rank. Eg. In wolf packs, relatives of the alpha male and female hold a higher rank than other wolves which may be stronger than them.
Subordinance/submissive
Individuals who display behaviours that show they are at the bottom of a social ranking eg. rolling over, displaying neck, averting eye contact.
Eusocial groups
Highly cooperative species where there is a division of labour into reproductive and non-reproductive groups. All individuals are strongly related eg. all bees come from a single queen.
Courtship displays
Behaviours aimed to facilitate attraction with the opposite sex for reproduction
Purposes of courtship
Attraction of opposite sex in same species
For animals requiring internal fertilisation, it makes the female more receptive rather than aggressive
Both sexes are fertile at the same time
Females can select the same mate
K-strategists
Heavy investment in few offspring, requiring more parental care and a longer maturation period.
R-Strategists
Many offspring are produced in one breeding season, and little to no parental care is given.
Monogamy
Males mate with one female in a breeding season
Why are some animals monogamous?
The energy required to incubate and feed the offspring is greater than the female can provide on her own, so male must ensure that his genes are passed on
Polygamy
Males mate with more than one female in a breeding season
Why are most mammals polygamous?
The female requires less energy to rear the offspring on her own. It is better for the male to increase his genes in the gene pool.
Polygynandry
A mating system where both males and females have multiple mating partners in a breeding season and young are cared for by several adults.
More parental care will increase the chance of survival of the offspring but will decrease the chance of survival for the parent
Viviparous: (of an animal) bringing forth live young which have developed inside the body of the parent, receiving food and protection
Oviparous: (of an animal) producing young by laying eggs which are hatched after the parent has laid them. Fertilisation is internal.
Altrical young are born helpless and blind
Precocial young are born developed and can walk and feed themselves
Importance of courtship behaviours
-Innate behaviour must be practised before the outcome to develop a strong pairbond
-Selection is crucial for reproductive success
-increases future recognition and reduces aggression and agonistic behaviour between courting couple in future matings