At high tide they are completely submerged by the ocean so experience similar abiotic factors (e.g. pH, salinity, temperature, etc. to the ocean ecosystem
At low tide they experience more extreme abiotic conditions (e.g. higher salinity and temperatures-only some organisms can tolerate these conditions
30% of the total energy available (75% of the gross productivity) is lost to the environment when organisms use energy produced from respiration for movement or body heat