Members of related populations that compete more with each other than with other species. Differences between members in form and behavior are related to differences in the ecological resources the species exploit
A species is a set of organisms sharing a common ancestor of monophyletic origin and similar evolutionary history. Members possess traits that combine them together (synapomorphy) and separate them from other species (autapomorphy)
The study of Alejandro et al., (2005) is titled "Polyphyly of Mussaenda Inferred from ITS and trnT-F Data and Its Implication for Generic Limits in Mussaendeae (Rubiaceae)"