Tall, has a woody stem (smaller woody plants = shrubs), monopodial growth (one trunk rather than many), apical dominance (grows upwards more quickly than branches grow outwards), secondary growth (trunk grows outward as well as upwards)
Families of trees
Arecaceae - date palm
Secondary thickening
A ring of dividing cells (meristem) just underneath the bark called the vascular cambium produces phloem on the outer side and xylem on the inner side
Sapwood = newer, outer xylem tissue (light-coloured), transporting water – lignified cells which are functionally dead
Heartwood = oldest, inner xylem, no transport, can rot away and not kill tree
Growth rings = seasonal growth
Monocot stems
Palm trees
Trees are water fountains
First trees evolved in the Northern hemisphere
390Ma (Middle Devonian Period)
Archaeopteris evolved in the Northern hemisphere
380Ma (Late Devonian Period)
Archaeopteris
Wood similar to conifers
Up to 8m tall, trunk diameter up to 1.5m
Still produced spores, not seeds
A progymnosperm
Formed vast, monodominant forests
Giant clubmosses (Lepidodendron) evolved in the Northern hemisphere
370-300Ma (Late Devonian to Upper Carboniferous)
Lepidodendron
Sometimes wood, lots of structural bark
10-35m tall, trunk diameter up to 1m
Produced spores, not seeds
A lycopsid (clubmoss)
Formed two-thirds of earliest tropical wetland forests
Full height in 10 years?
Calamites and Medullosa evolved in the Northern hemisphere
360-252Ma (Carboniferous-Permian)
Calamites
Woody, lots of structural bark
Up to 18m tall, trunk diameter up to 1m
Produced spores, not seeds
A sphenopsid (horsetail)
Formed thickets between lycopsid trees, rather than understorey
Conifers in all biomes
Early Jurassic (206-180Ma)
Angiosperm revolution around 100Ma, with more than 400,000 species of flowering plant today (compare with 750 species of seed plants), worldwide dominance by end-Cretaceous