The study of interactions among organisms and between the organism and its physical (abiotic) environment
Levels of biological organization in ecology
Organisms
Populations
Communities
Biomes
Physiological ecology
The study of adaptation of an organism to environments in terms of survival and reproduction
The rotation of earth and the tilt of its axis cause annual variations in temperature & seasons. Major biomes (desert, rain forest, tundra etc.) are formed due to these variations & precipitation (rain & snow)
Regional and local variations within a biome lead to the formation of different habitats
Life exists even in extreme & harsh habitats
Niche
The distinct role and position of an organism in its environment
Abiotic factors
Temperature
Water
Light
Soil
Eurythermal
Organisms that can tolerate a wide range of temperatures
Stenothermal
Organisms that can tolerate only a narrow range of temperatures
Euryhaline
Organisms that tolerate a wide range of salinities
Stenohaline
Organisms that tolerate only a narrow range of salinity
Regulate
The maintenance of homeostasis by physiological & behavioural means
Conform
Organisms that cannot maintain a constant internal environment, their body temperature or osmotic concentration change with the surrounding conditions
Migrate
Animals that move away temporarily from stressful habitat to a more hospitable area and return when stressful period is over
Suspend
Organisms that reduce their metabolic activity to survive unfavourable conditions
Adaptation
The morphological, physiological & behavioural attribute that enables an organism to survive and reproduce in its habitat
Adaptations of kangaroo rat in North American deserts
Internal fat oxidation gives water as byproduct if there is no external source of water
Ability to concentrate urine so that minimal volume of water is used to remove excretory products
Adaptations of desert plants
Presence of thick cuticle on leaf surfaces
Sunken stomata minimise water loss due to transpiration
CAM photosynthetic pathway enables their stomata to remain closed during day time
Desert plants like Opuntia have no leaves (they are reduced to spines), photosynthesis is done by stems
Adaptations of mammals
Mammals from colder climates have shorter ears and limbs to reduce heat loss (Allen's Rule)
Aquatic mammals like seals have a thick layer of fat (blubber) below their skin that acts as an insulator and reduces loss of body heat
Physiological and biochemical adaptations
Archaebacteria are found in hot springs & deep-sea hydrothermal vents where temperature is >100??C
Many marine invertebrates & fishes live at great depths in the ocean where the pressure is >100 times the normal atmospheric pressure
At a high-altitude place (>3,500 m) the body compensates low O2 availability by increasing RBC & breathing rate and decreasing the binding capacity of hemoglobin
Behavioural adaptations
Desert lizards bask in the sun and absorb heat when their body temperature is low, but move into shade when the ambient temperature starts increasing
Some species burrow into the soil to hide and escape from the above-ground heat
Population
A group of individuals of same species that live in a given geographical area, share or compete for similar resources and potentially reproduce
Birth rate
Per capita births
Death rate
Per capita deaths
Sex ratio
The proportion of males and females in a population
Age pyramid
The structure obtained when the age distribution (% individuals of a given age or age group) is plotted for the population
Population size or population density (N)
The number of individuals of a species per unit area or volume
Natality (B)
The number of births in a population during a given period
Mortality (D)
The number of deaths in a population during a given period
Immigration (I)
The number of individuals of the same species that have come into the habitat from elsewhere during a given time period
Emigration (E)
The number of individuals of the population who left the habitat and gone elsewhere during a given time period
Population growth
Nt+1 = Nt + [(B + I) - (D + E)]
Exponential growth
If resources are unlimited, each species shows its full innate potential to grow in number, and the population grows in an exponential or geometric fashion
Intrinsic rate of natural increase (r)
An important parameter for assessing impacts of any biotic or abiotic factor on population growth
When colonizing habitat, immigration may be more significant to population growth than birth rates
Exponential growth
Resources (food & space) are essential for the unimpeded population growth
If resources are unlimited, each species shows its full innate potential to grow in number. Then the population grows in an exponential or geometric fashion
If population size = N, birth rates (per capita births) = b and death rates (per capita deaths) = d, then the increase or decrease in N during a unit time period t (dN/dt) will be dN/dt = (b – d) × N
Let (b–d) = r, then dN/dt = rN
The r ('intrinsic rate of natural increase') is an important parameter for assessing impacts of any biotic or abiotic factor on population growth
Logistic growth
There is no population in nature having unlimited resources for exponential growth. This leads to competition among individuals for limited resources
Eventually, the 'fittest' individuals survive and reproduce
In nature, a given habitat has enough resources to support a maximum possible number, beyond which no further growth is possible. It is called carrying capacity (K)
A population with limited resources shows initially a lag phase, phases of acceleration & deceleration and finally an asymptote. This type of population growth is called Verhulst-Pearl Logistic Growth. It is described by following equation: N = K / (1 + (K-N0)/N0 * e^(-rt))
Life History Variation
Populations evolve to maximise their reproductive fitness or Darwinian fitness (high r value)
Some organisms breed only once in their lifetime (Pacific salmon fish, bamboo) while others breed many times (most birds and mammals)
Some produce a large number of small-sized offspring (Oysters, pelagic fishes) while others produce a small number of large-sized offspring (birds, mammals)
These facts indicate that life history traits of organisms have evolved due to limited abiotic and biotic components of the habitat